Zoophycos

Zoophycos may be a multi-functional feeding garden model of trace maker according to the fillings and pyrite framboids in light-and dark-grey minor lamellae of burrow (Gong et al. 2007 (Gong et al ...

Zoophycos. Zoophycos is one of the most complex and enigmatic trace fossils recorded in marine strata from Cambrian to Quaternary worldwide, which is invaluable for the study of Phanerozoic development of organism-environment interactions. Here we address and demonstrate the macroevolution of Phanerozoic Zoophycos by assembling 448 points in constructing the Phanerozoic Zoophycos database based on 291 ...

The Zoophycos Ichnofacies is characterized by complex fodinichnia (Zoophycos, and sometimes other deep traces such as Thalassinoides) in tiered arrangements. (after Benton & Harper, 1997) The ichnofacies occurs in a range of water depths between the abyssal zone and the shallow continental shelf, in normal background conditions of sedimentation.

An ethological interpretation of Zoophycos based on Permian records from south China and Southeastern AustraliaZoophycos ispp. domina en fangolitas de talud con deslizamientos sinsedimentarios. Concordantemente con una marcada tendencia de enfriamiento, el Eoceno Medio superior-Moreover, the Zoophycos ichnogenus has been studied and debated for more than a century without reaching a consensus on its ethologies and probable trace makers, however, it has been considered as ...Zoophycos from both regions is composed of a marginal tube and a tongue-like spreiten complex, the latter itself consisting of primary lamellae in planar view and backfill structures (dark and ...Zoophycos is most common in sediments deposited during glacial times and when the sedimentation rate was intermediate and primary production was high and seasonal. Occurrences of Zoophycos elsewhere support a similar relationship with seasonal organic-matter deposition. This is particularly significant considering that seasonality of organic ...Firstly, the fractal topography in Zoophycos was established by introducing virtual tertiary lamellae. Next, a fractal dimension ( D ) model was proposed and the spatio-temporal evolution trend of the complexity of the trace-making behavior was established.

Echinospira isp., a relatively small, lobate, trace fossil of the Zoophycos group, occurs in Miocene flysch deposits of the Cingöz Formation of southern Turkey. It displays a burrow system which co...As noted from terrestrial occurrences (Ekdale 1992 ), Zoophycos is a deep tier trace fossil produced up to a metre beneath the sediment-water interface. An age difference of 9 ka has been found between the sediment within the Zoophycos spreite and the surrounding sediment (Leuschner et al. 2002 ). Zoophycos isp (Fig. 6 E and F). occurs as epichnia in 10-15 cm-thick, yellow, fine-grained, sandstone of the Tiaret Marl Formation in The Merzeghellal locality. Zoophycos isp. is a spreiten structure consisting of a U-shaped burrow and lamellae spiraling around a central tube burrow. The individual lamellae are 150-200 mm long and 4-6 mm ...C. Somite. D. Bracing. zoophycos pronunciation - How to properly say zoophycos. Listen to the audio pronunciation in several English accents.The Zoophycos ichnofacies (Fig. 19.13b) is characterized by complex fodinichnia like Zoophycos, and it may contain other deep traces such as Thalassinoides in tiered arrangements. The ichnofacies occurs in a range of water depths between the abyssal zone and the shallow continental shelf, and may be associated with low levels of oxygen. This ...

Ichnology is the study of trace fossils, the record of microbe, plant, and animal behavior preserved in rocks. Ichnology (not to be confused with Ichthyology—the study of fish) is a major subdiscipline in the field of paleontology , which is concerned with the study of ancient life at large. Ichnology lies at the crossroads of paleontology ...Ichnofacies can provide information about water depth, salinity, turbidity and energy. In general, traces found in shallower water are vertical, those in deeper water are more horizontal and patterned. [1] This is partly because of the relative abundance of suspended food particles, such as plankton, in the shallower waters of the photic zone ...Zoophycos isp. showing (B) the wider lobe in its distal part, AG-51(115) ... Zoophycos ichnofacies. Palaios 6, 410–425. Miller, M.F., 2003. Styles of ...Zoophycos is one of the most widespread but still enigmatic trace fossils, especially with regard to morphology and ethology surrounding its 3-D helically coiled spreite structures. The genus Zoophycos is systematically revised based on the systematic researches of the literature with detailed description and plate from 1855 to 2011.Zoophycos was only found in two cores taken at more than 2 km water depth on the slope of the Lomonosov Ridge, but was absent in cores obtained at shallower depth, confirming earlier observations of the trace maker's bathymetric preferences. The two cores containing Zoophycos are characterized by quiet sedimentation and slightly enhanced food ...Palaeophycus, Zoophycos, Chondrites, Lorenzinia, and Spirorhaphe. Ophiomorpha It has an irregular tube shape, filled with agglutinated sediment lumps/pellets on its walls, the texture

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The Zoophycos ichnofacies ideally is found in circalittoral to bathyal, quiet-water marine muds or muddy sands, below the storm wave base, particularly in shelfal to slope areas (MacEachern et al., 2007, MacEachern et al., 2012; Buatois and Mángano, 2011, Fig. 12). Settings are generally free of turbidity flows and subject to oxygen deficiencies.The trace fossil Zoophycos is abundant in transgressive, shallow marine carbonates in the Middle Permian (Wordian) Khuff Formation of the Huqf-Haushi Uplift of Interior Oman. It often occurs as part of a complex (compound) trace fossil that comprises two integrated elements: (i) irregular galleries with straight to gently curved tunnels and interconnected shafts, and (ii) simple planar to ...Zoophycos is a deep tier structure with several ethological interpretations, but consensus favors the interpretation of cache behavior developed by vermiform animals 27,28.Zoophycos is a complex and enigmatic ichnofossil that has challenged generations of geologists and ichnologists for over a century. Till now, its producer or producers remain elusive. This ichnofossil is cosmopolitan and it occurs in diverse marine sedimentary facies ranging from Palaeozoic to Holocene.Zoophycos from both regions is composed of a marginal tube and a tongue-like spreiten complex, the latter itself consisting of primary lamellae in planar view and backfill structures (dark and ...Icnoespecie:Zoophycos isp. Descripción: estructura icnológica formada por superficies estriadas y lobuladas que se disponen helicoidalmente alrededor de una galería central que no e visible. Posible artrópodos o anélidos formaron la estrucutra en un ambiente de talud continental/llanura abisal, con poca oxigenación y una sedimentación de baja velocidad en un ambiente de muy poca energía.

Palaeozoic Zoophycos occurs in marine margin, shal- low water, and deep water deposits (Chamberlain 1971,. Osgood & Szmuc 1972, Miller 1991). Mesozoic Zoophy-.The Zoophycos ichnofacies ideally is found in circalittoral to bathyal, quiet-water marine muds or muddy sands, below the storm wave base, particularly in shelfal to slope areas (MacEachern et al., 2007, MacEachern et al., 2012; Buatois and Mángano, 2011, Fig. 12). Settings are generally free of turbidity flows and subject to oxygen deficiencies.Distribusi geografis fosil memungkinkan ahli geologi untuk mencocokkan komposisi batuan dari bagian lain dunia. 6 Jenis-Jenis Fosil: Pengertian, Persebaran, Proses, Syarat, Tipe dan Pemanfaatan - Fosil …May 1, 2022 · As such, lobed Zoophycos are rare in the Palaeozoic but common in post-Palaeozoic times. This study reports an unusually lobate Zoophycos from the early Permian Tahkandit Limestone (Brabb and Grant, 1971) of the Yukon–Charley Rivers National Preserve, east-central Alaska. These traces consist of numerous vertically oriented whorls and ... Variations of the trace fossil Zoophycos with respect to paleoenvironment and sequence stratigraphy in the Mississippian Mobarak Formation, northern Iran Bayet-Goll, Aram Daraei, MehdiZoophycos is widely distributed in the marine strata of the Middle Permian Ruteh For- mation in the Alborz Mountain, Iran. The investigation of the Zoophycos , along with environmental variables ...Abstract. Zoophycos is a complex three dimensional trace fossil that is abundant in deep ocean sediments worldwide, but has not been described previously from Cenozoic …

kotake, n, paleoecology of the zoophycos producers, lethaia 22: 327 (1989). google scholar. kyte, f.t., accretion rate of extraterrestrial matter - iridium deposited 33 to 67 million years ago, science 232: 1225 (1986). crossref. pubmed. isi. google scholar. lowrie, w, earth planet sc lett 98: 303 (1990). google scholar. lowrie, w, 100 million years of …

Terdapat fosil jejak berupa Nereites yang merupakan penciri dari lingkungan pengendapan laut dalam (deep marine) atau bathyal. Sedangkan fosil jejak berupa Zoophycos yang merupakan bentuk transisi, dapat sebagai penciri lingkungan pengendapan offshore shelf sampai deep marine (abysal-bathyal). III.2 LOKASI PENELITIAN II (STOP SITE II)kotake, n, paleoecology of the zoophycos producers, lethaia 22: 327 (1989). google scholar. kyte, f.t., accretion rate of extraterrestrial matter - iridium deposited 33 to 67 million years ago, science 232: 1225 (1986). crossref. pubmed. isi. google scholar. lowrie, w, earth planet sc lett 98: 303 (1990). google scholar. lowrie, w, 100 million years of …Zoophycos is widely distributed in the marine strata of the Middle Permian Ruteh For- mation in the Alborz Mountain, Iran. The investigation of the Zoophycos , along with environmental variables ...Zoophycos-group burrows are prevalent elements of the post-Cambrian trace fossil record.Here we report the oldest specimens of Zoophycos from Lower Cambrian strata of the Lower Member Wood Canyon Formation in southeastern California. In addition to these being the oldest examples of this well-known trace fossil, the discovery of these specimens also reveals the presence of deposit feeding ...The Zoophycos ichnofacies is reported for the first time from the Lower Jurassic (Toarcian) deposits of Algeria. The trace fossil-bearing strata in the Ksour Mountains, northwestern Algeria, are an alternation of fine-grained limestones and marls. The assemblage is characterised by its low diversity, consisting of eight ichnospecies (Chondrites intricatus, C.Ichnofacies can provide information about water depth, salinity, turbidity and energy. In general, traces found in shallower water are vertical, those in deeper water are more horizontal and patterned. [1] This is partly because of the relative abundance of suspended food particles, such as plankton, in the shallower waters of the photic zone ...The producer of the lobed Zoophycos of the Yukon River is interpreted as a deposit-feeder that used sensory-driven, directed search for locating heterogeneously distributed trophic resources. The Zoophycos producer filled its burrow with Coprolus-like fecal pellets, possibly complementing deposit feeding with microbial gardening and/or food ...Zoophycos comprises a diverse group of spreiten structures. There are only a few published reports of Zoophycos from New Mexico and these are primarily from Carboniferous strata.Aug 1, 2020 · Multina–Zoophycos and multi-tier Zoophycos ichnofabrics, representing vertical migration of benthic faunas in pace with either distal event deposition or steadily accreted sea floor. The colonisation window in the arc-related setting is closely related to sedimentation rates and frequency of depositional events, determining the completeness ...

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Zoophycos is a complex fodinichnial structure (Bromley and Hanken, 2003). The Palaeozoic Zoophycos are documented from shallow to deep-sea deposits, the Mesozoic forms in the deeper shelf and occurring in deeper waters since the Middle Jurassic age ( Olivero, 2003 ), while the Cenozoic Zoophycos are recorded mostly in deep-sea facies (e.g ...Title : ( Occurrence of Zoophycos in the Ruteh Formation, Middle Permian (Guadalupian), Central Alborz, Iran: palaeoenvironmental and sequence stratigraphy ...Zoophycos is a characteristic member of the Zoophycos ichnofacies (Pemberton et al., 2001) and it represents the offshore environment, where it inhabits deep levels in the sediment (Solanki et al ...A 425 ka long record from piston core GIK17925-3 taken in the northeastern South China Sea was used to study the environmental conditions controlling the distribution of the Zoophycos trace fossil. The 12 m long core offers a unique opportunity to study the response of the Zoophycos-producing animal to environmental variations over four glacial-interglacial cycles.Zoophycos gibi.), çakmaktaìlarl içinde gözlenmesi dikkat çekicidir Bitki fosillerinden ise nadir de Olsa çok iyi örnekler çakmakta§larl içinde bulunmu#ur. Kuzey Avrupa'da Geç Kretase ya§ll sedimanter kayalar içindeki çakmaktay bantlaflnda denize çok yakln bir §ekilde yaìamlss kozalakll agaçlara ait Cryptomeriopsis eluvialis türü bitki fosilleriThalassinoides paradoxicus (Woodward, 1830) is a prevalent and frequently reported ichnotaxon, but given its early erection and subsequent revisions, it now is poorly understood and lumps together three different kinds of trace fossils having distinct characteristics. A clear definition (i.e., diagnosis) of each individual component is crucial for ichnotaxonomic stability and the application ...El icnogénero Zoophycos y su valor sedimentológico y paleoambientalOct 9, 2015 · Palaeozoic Zoophycos caudagalli and Mesozoic–Cenozoic Zoophycos insignis both started with an initial U-tube or J-tube, and developed into sub-circular or lobate patterns. Feeding, gardening and farming in this proposed phylogenetic sequence are more a continuous track of improving efficiency rather than simple, discrete behavioral patterns. Zoophycos is a complex fodinichnial structure (Bromley and Hanken, 2003). The Palaeozoic Zoophycos are documented from shallow to deep-sea deposits, the Mesozoic forms in the deeper shelf and ... ….

The first Zoophycos-bearing level (Zl 1) is characterised by an overall high bioturbation intensity reflecting a very high oxygenation rate and nutrient supply, allowing the development of large ...Zoophycos from both regions is composed of a marginal tube and a tongue-like spreiten complex, the latter itself consisting of primary lamellae in planar view and backfill structures (dark and ...The substrate-controlled Glossifungites ichnofacies is a firmground suite of trace fossils that commonly demarcates erosional discontinuities in sedimentary successions. The Glossifungites ichnofacies typically is characterized by biogenic structures that are vertical to subvertical, sharp-walled, unlined and commonly passively infilled. The structures hitherto have been restricted to ...Firstly, the fractal topography in Zoophycos was established by introducing virtual tertiary lamellae. Next, a fractal dimension (D) model was proposed and the spatio-temporal evolution trend of the complexity of the trace-making behavior was established. In application, the correlation of D to the paleo-water depth and substrate consistency of ...Request PDF | On Dec 1, 2015, David Gallego-Torres and others published Pyrite framboid size distribution as a record for relative variations in sedimentation rate: An example on the Toarcian ...However, the trace fossil assemblage (Planolites, Chondrites, Zoophycos, Thalassinoides, and Teichichnus; Fig. 3; Table DR1) and consistent presence of organic benthic foraminiferal linings and Mn (Fig. 2; Fig. DR1) throughout the Site 959 study section indicate at least occasional seafloor oxygenation. Combined, these lines of evidence ...Zoophycos - simple planar form conditions (Brasier, 1996). morphology (drawn after photo of the Upper Lower diversity within benthic foraminiferal Bathonian Zoophycos sample from Alpes de assemblage as well as small size and pre- Haute Provance, SE France) dominance of flattened morphologies has been frequently observed in deposits that were ...Zoophycos are spiral, concentric, lobate structures that are filled with mud or organic matter; they demonstrate deep water, low sedimentation rate, and high biotic diversity.Zoophycos is a complex burrow with stacked tiers, in which the tiers are made of spreiten-adorned lobes that radiate outward from a central tube in a downward spiral to form a shape that looks something like a Christmas tree in three-dimensions. Each lobe contains a peripheral tube, with spreiten loops between the tubes. The spreite in map view have "rooster tail" patterns that spiral out from ...Zoophycos is a complex fodinichnial structure (Bromley and Hanken, 2003). The Palaeozoic Zoophycos are documented from shallow to deep-sea deposits, the Mesozoic forms in the deeper shelf and occurring in deeper waters since the Middle Jurassic age ( Olivero, 2003 ), while the Cenozoic Zoophycos are recorded mostly in deep-sea facies (e.g ... Zoophycos, Zoophycos is one of the most complex and enigmatic trace fossils recorded in marine strata from Cambrian to Quaternary worldwide, which is invaluable for the study of Phanerozoic …, , Nov 1, 2013 · The Zoophycos producer filled its burrow with Coprolus-like fecal pellets, possibly complementing deposit feeding with microbial gardening and/or food caching. Data presented here provide useful insight into the morphological evolution and bathymetric distribution of Zoophycos , suggesting two ‘Golden Ages’ for lobed Zoophycos: (1 ... , Pa (Palaeophycus); Th (Thalassinoides); Zo (Zoophycos) Full size image. Conclusions. 1. The digital image treatment described here has been applied to core sediments from the IODP Expedition 339, Site U1385, allowing modifications in certain image features such as contrast, brightness, vibrance, saturation, exposure, lightness, …, Since the Jurassic, chalks and related carbonates have accumulated via multiple processes, under a broad range of water depths and in a wide array of marine environments. Trace-fossil suites in these deposits are highly variable, representing as many as six ichnofacies. Variability among Zoophycos Ichnofacies suites reflect variations in water ..., The Zoophycos ichnofacies ideally is found in circalittoral to bathyal, quiet-water marine muds or muddy sands, below the storm wave base, particularly in shelfal to slope areas (MacEachern et al., 2007, MacEachern et al., 2012; Buatois and Mángano, 2011, Fig. 12). Settings are generally free of turbidity flows and subject to oxygen deficiencies., Last Updated: 16 Aug 2023. PDF | Zoophycos is one of the most complex and enigmatic trace fossils recorded in marine strata from Cambrian to Quaternary worldwide, which …, 1. Introduction. Zoophycos is among the most distinct trace fossils. As defined by Häntzschel (Reference Häntzschel and Teichert 1975), Zoophycos is a planar spreite marking successive, former positions of the burrow, present either as simple lobes or as ornate spiral structures, enclosed by a marginal burrow which may not always be evident (either not developed or not preserved)., Zoophycos isp. is an enigmatic trace that has been related to different organisms, including sipunculids, echiurians, polychaete and worms, and assigned to various ethologic models such as strip-mine, detritus feeding, gardening, refuse-dump, and cache models ..., Zoophycos Massalongo, 1855 DESCRIPTION: Spreiten structures consisting of numerous distinct U-shaped, protrusive lobes of variable length, bordered by a marginal tunnel (i.e., causative burrow). Lobes coil spirally around a central axis, which may be a spreiten whorl structure. BEHAVIOR(S): Fodinichina; feeding burrows of deposit feeders. ENVIRONMENTAL SETTINGS: Shallow to deep marine, The modern Zoophycos found in deep-sea sediments from the Portuguese continental margin differs markedly from most Zoophycos described from the fossil record. A detailed X-ray radiograph analysis revealed that the trace consists of an upwards constructed, helically coiled spreite that is connected to the sediment surface through a single shaft., The Zoophycos ichnofacies is characteristic of such oxygen-deficient settings as restricted littoral to bathyal quiet-water environments (Fig. 14). This ichnofacies is typical in offshore environments below maximum storm wave base free of turbidity deposition and strong bottom currents. , INTRODUCTION. Zoophycos is a trace fossil whose exact organisation and significance have been discussed for long. According to the modern interpretations, it is a complex spreiten structure with a tremendous morphological variability (Bromley, 1996).Numerous descriptions have appeared since its first description in 1855 by Massalongo (see Olivero, Chapter 13), and this has led to a general ..., The complex and enigmatic trace fossil Zoophycos is recorded worldwide, from Cambrian to Holocene. The possible evolution of the ichnofossil has already been proposed by some authors, but their works concerned the very large group of trace fossils related to Zoophycos (structures that could be assigned potentially to several ichnogenera) recorded in several depositional basins., The trace fossil Zoophycos was enriched in the lower part of the Guadalupian Maokou Formation of. Permian in Laibin, Guangxi of South China, ..., This unit exhibits a second positive CIE (δ 13 C org increases by 2.5‰ and δ 13 C carb increases by 1.5‰), significantly more negative δ 18 O values relative to the pre-T-OAE interval, and a low diversity of trace fossils dominated by Chondrites and extremely thin Zoophycos. Sponge spicules re-appear in the uppermost 6.5 m-thick interval of the …, May 21, 2019 · Detailed ichnological and petrographic analyses were conducted on two biostratigraphically-constrained intervals (11.9-11.3 Ma and 10.7-9.4 Ma) from the Middle-Upper Miocene stratigraphy of the Great Bahama Bank’s leeward margin to assess spatiotemporal trends in paleoenvironmental conditions and reservoir properties in carbonate slope deposits. Six ichnofacies, varied in expression, are ... , Zoophycos ichnofacies was associated with chalky calcilutite sediment, interpreted as calcareous muddy contourites deposited during low-energy conditions (Hüneke et al., 2020). Increased energy in the environment led to a shift toward the first sandy contouritic facies and the establishment of the distal Cruziana ichnofacies with predominant ... , Zoophycos trace fossil in the Mississippian of Ohio, USA. This rock sample is from the upper Logan Formation of eastern Ohio. The curving grooves are part of a trace fossil called Zoophycos, a distinctive feeding trace with an overall spiral morphology (for example, see: ..., De, 1998, De, 1999b studied 12 ichnotaxa such as Chondrites, Cylindrichnus, Ophiomorpha, Planolites, Psilonichnus, Rosselia, Skolithos, Teichichnus, Thalassinoides, Terebellina, Zoophycos and spiral gastropod burrows from the Barakar Formation in this basin. These ichnofossils are reported from three outcrop sections (one in Angul-Kosala …, Zoophycos is one of the most complex and enigmatic trace fossils recorded in marine strata from Cambrian to Quaternary worldwide, which is invaluable for the study of Phanerozoic development of organism-environment interactions. Here we address and demonstrate the macroevolution of Phanerozoic Zoophycos by assembling 448 points in constructing the Phanerozoic Zoophycos database based on 291 ..., The different ethological modes of Zoophycos producer exerts ambiguity to the use of this trace fossil (in conjunction with sedimentological data) as an environmental indicator, especially in evaluating energy level, substrate consistency, and organic matter distribution or nutrient supply. The problem is even more sophisticate within the sequence stratigraphic framework., Zoophycos is found off NW Africa in water depths > 2,000 m, in sediments with C org contents of 0.3-1.8%. It is possible to recognize in the core sections all of the morphological spreiten types of Zoophycos which were described from ancient sediments. All of these spreiten are formed with two modifications classified as "U-type" and "J ..., The Zoophycos ichnocoenosis was produced during a further later but relatively long stage after the emplacement of the turbidites, indicating a much more reducing bottom-water condition., The Cambrian explosion represents one of the most dramatic episodes in the history of the biosphere. Following a period during the Ediacaran when biotas were dominated by soft-bodied organisms, some of which seemingly unrelated with modern metazoans (Seilacher 1989; Narbonne 2005), the fossil record changed rapidly as …, This specimen shows clearly that it is Zoophycos in its earlier morphotype (although it hasn't a comparative scale) when it was smaller than the later ones in the evolution process , similar to the specimens from Zhang et al., 2015. excerpt from L. Zhang et al. 2015. Zoophycos macroevolution since 541 Ma. Scientific Reports, 5: 1-10, Zoophycos is one of the most complex and enigmatic trace fossils recorded in marine strata from Cambrian to Quaternary worldwide, which is invaluable for the study of Phanerozoic development of organism-environment interactions. Here we address and demonstrate the macroevolution of Phanerozoic Zoophycos by assembling 448 points in constructing the Phanerozoic Zoophycos database based on 291 ... , Zoophycos is one of the most complex and enigmatic trace fossils recorded in marine strata from Cambrian to Quaternary worldwide, which is invaluable for the study of Phanerozoic development of organism-environment interactions. Here we address and demonstrate the macroevolution of Phanerozoic Zoophycos by assembling 448 points in constructing the Phanerozoic Zoophycos database based on 291 ..., , Media in category "Zoophycos". The following 7 files are in this category, out of 7 total. Zoophycos fossil burrow (Borden Formation, Lower Mississippian; Portsmouth Bypass roadcut, Ohio, USA) (48118331742).jpg 3,964 × 2,752; 7.98 MB. Zoophycos mcr1.JPG 3,456 × 2,304; 6.37 MB. , Four Zoophycos morphotypes are recognized at Gubbio showing variations of major and minor lamellae, apex, lobes and whorls: the Cretaceous-Eocene cone-shaped type 1, the Upper Eocene-Middle ..., Thalassinoides paradoxicus (Woodward, 1830) is a prevalent and frequently reported ichnotaxon, but given its early erection and subsequent revisions, it now is poorly understood and lumps together three different kinds of trace fossils having distinct characteristics. A clear definition (i.e., diagnosis) of each individual component is crucial for ichnotaxonomic stability and the application ..., The feeding mechanism of Zoophycos producers demonstrates non-selection by particle size or quality. The characteristic morphology and mode of Occurrence of Zoophycos and the presence of pellets with pyroclastic grains imply that the Zoophycos producers were surface deposit-feeders. They dwelt in a burrow within the sediment and ingested the grains with nutrient detritus on the sea floor ...